Adaptation in Stochastic Environments by Jin Yoshimura, Colin W. Clark

By Jin Yoshimura, Colin W. Clark

The classical conception of average choice, as constructed through Fisher, Haldane, and 'Wright, and their fans, is in a feeling a statistical idea. normally the classical conception assumes that the underlying atmosphere within which evolution transpires is either consistent and reliable - the idea is during this experience deterministic. actually, nevertheless, nature is sort of consistently altering and volatile. we don't but own an entire conception of traditional choice in stochastic environ­ ments. probably it's been inspiration that any such conception is unimportant, or that it might be too tough. Our personal view is that the time is now ripe for the advance of a probabilistic conception of typical choice. the current quantity is an try and supply an ordinary advent to this probabilistic concept. each one writer used to be requested to con­ tribute an easy, easy advent to his or her forte, together with full of life discussions and hypothesis. we are hoping that the booklet contributes additional to the certainty of the jobs of "Chance and Necessity" (Monod 1971) as built-in parts of model in nature.

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1976. Life history tactics: A review of the ideas, Q. Rev. BioI. 51:3-47. C. 1992. The evolution of life histories. Oxford University Press. D. 1989. An uncertain life: demography in random environments, Theoret. Pop. Biol. 35:227-294. Yoshimura, J. W. Clark. 1991. Individual adaptations in stochastic envirnments, Evol. Ecol. 5:173-192. 6) DENSITY DEPENDENT LIFE HISTORY EVOLUTION IN FLUCTUATING ENVIRONMENTS Eva Kisdi Population Biology Group Department of Genetics, Eotvos University Budapest, Muzeum krt.

Ecol. 5:173-192. 6) DENSITY DEPENDENT LIFE HISTORY EVOLUTION IN FLUCTUATING ENVIRONMENTS Eva Kisdi Population Biology Group Department of Genetics, Eotvos University Budapest, Muzeum krt. 4/ A, Hungary, H-I088 and Geza Meszena Population Biology Group Department of Atomic Physics, Eotvos University Budapest, Puskin u. 5-7, Hungary, H-I088 Abstract. Environmental fluctuation may not only alter the life history optimization problem but also query optimization itself. Under density regulation annual growth rate is influenced by the direct effect of fluctuation as well as by an indirect effect due to fluctuating population density.

Competition for a single resource in a stable environment. Strategy 3 with the highest equilibrium density (Ka) can initially increase in the equilibrium population of strategy lor strategy 2, since its annual growth rate exceeds 1 at density N = Kl as well as at N = K 2 . Invasion of strategy 3 increases the density, hence the annual growth rate of the former strategy lor 2 becomes lower than 1: strategy 3 spreads and the former strategy will be excluded from the population. The population reaches its evolutionary stable equilibrium at N = K a, when strategy 3 is established.

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